The world has bid farewell to a remarkable creature whose life bridged three centuries. The San Diego Zoo recently mourned the passing of Gramma, a Galápagos tortoise estimated to be 141 years old—a gentle giant who outlived two world wars, more than 20 U.S. presidents, and two global pandemics. Born around 1884 in the Galápagos Islands, Gramma arrived at the zoo between 1928 and 1931 as part of an early conservation initiative. Affectionately dubbed the "Queen of the Zoo" by her caregivers for her sweet, shy demeanor, she captivated generations of visitors, evolving from a figure in vintage black-and-white photos to a modern social media icon. In November 2025, zoo staff made the compassionate choice to humanely euthanize her due to progressive age-related bone degeneration. As an enduring symbol of her endangered species, Gramma inspired millions to support reptile conservation efforts worldwide. To celebrate her legacy, the zoo suggests honoring her with one of her favorite meals: a fresh salad featuring romaine lettuce and cactus fruit. Though Gramma is no longer with us, her extraordinary longevity and quiet presence continue to underscore the San Diego Zoo's vital role in protecting Galápagos tortoises.